Abstract: Explaining how cooperation evolves is one of the primary 
research projects in evolutionary biology and philosophy of biology. 
Such work typically focus on the maintenance of cooperation in the 
face of potential free-riders. While this is no doubt important, 
explaining how living in a group "generates benefits" for individuals 
over solitary living is also necessary for a full account of the 
evolution of cooperation. In this paper, I consider this "second 
problem of cooperation" from a new perspective focusing on social 
learning in non-human animals and its impact on the evolution of 
cooperation.